So I bypassed Crusher shot by using Ekoro's arrows to break blocks. When entering Area 3, there is a set of blocks that can be destroyed from the side when going down, but have to be shot from below when going up. Since Ekoro can only fire arrows horizontally in overworld, I can't find a way to break these block.

Did I just softlocked myself? Here is a screenshot. Only all upgrades from Area 1 and Area 3 have been collected.

https://steamcommunity.com/sharedfiles/filedetails/?id=3523569050